I cbf going through 3 pages of discussion, so sorry if this has already been posted:
I was under the impression that planes lift off because of the reaction force acting up by air on plane (mainly the wings, it's somehow related to Bernoulli, but I forgot exactly how, but the design of the wings gives a pressure difference hence 'sucks' the plane upwards). In this case, if we want to model this, the force we're looking at is related to the relative velocity with air.
The engine generate a net turbulence towards the back of the plane [takes in air from the front, release the products of combustion as hot gases at the back which expands :. propel plane forward]. Hence, the plane has a driving force forwards [and net acceleration forwards].
The wheels are in contact with the conveyor belt, but if we ignore the friction between the wheel and its axle (and assume the entire driving and breaking forces are all supplied by the engine), the wheels are just rolling freely. They are rotating because of the static friction between the surface (conveyor belt) and the surface of the wheel (try a pen rolling across a table).
That is, the wheels in this case does not transfer any forces from the conveyor belt to the plane. Hence, if the plane is accelerating at +a due to the thrust force, and there is no retarding forces acting on it from the conveyor belt, then its motion relative to air is the same as before (assuming the conveyor belt doesn't generate too huge a turbulence against what the plane is trying to achieve). Therefore, it takes off however it was taking off before, just, with the wheels spinning madly.
Yes, the plane does take off. But I wouldn't want to be the wheels.
[if it was say, a car going up a jump ramp, and the jump ramp was a conveyor belt, then no, the car wouldn't be able to do the jump (wouldn't be able to go up the ramp at all). In this case, the driving force is supplied by the friction between the wheel and the road surface.]
I might have made some critical errors though. I don't have enough time to think this over again in terms of energy, but I have a feeling that the extra rotational energy of the wheels come from the energy of the conveyor belt, which has a driving force, and things should balance. But again taking off can be considered as a 'collision' between Earth and the plane, and there need to be consideration on how this 'collision' takes place in terms of momentum as well... so...